diff --git a/README.md b/README.md
index 60c3836..7c310ed 100644
--- a/README.md
+++ b/README.md
@@ -136,6 +136,11 @@ x = []  # type: list[int]
 To change single colors, override the groups in your vimrc after your
 colorscheme (with the palette on, a later `:colorscheme` re-applies it):
 
+Brackets, commas and colons inside annotations are not colored, like the rest
+of Python's punctuation. To color them, link `pythonTypeBracket`,
+`pythonTypeComma`, `pythonTypeColon`, `pythonParams` or `pythonDefColon`, e.g.
+`hi link pythonTypeBracket Delimiter`.
+
 ```vim
 " Example: Make typing types cyan instead of orange
 hi pythonTypingType     ctermfg=44  guifg=#00d7d7
diff --git a/doc/python-syntax-enhanced.txt b/doc/python-syntax-enhanced.txt
index 7a097a1..cd90e9f 100644
--- a/doc/python-syntax-enhanced.txt
+++ b/doc/python-syntax-enhanced.txt
@@ -173,7 +173,9 @@ Type Annotation Groups:~
     pythonTypeName          CapWords / aliases / TypeVars in annotations
     pythonTypeDotted        Dotted type paths (collections.abc.Sequence)
     pythonTypeUnion         Union operator `|` (inside type regions only)
-    pythonTypeBracket       Generic brackets `[` `]` and type parentheses
+    pythonTypeBracket       Brackets `[` `]` and parentheses in a type (*)
+    pythonTypeComma         Commas in a type (*)
+    pythonTypeColon         Colon before an annotation (*)
     pythonTypeComment       Type comments `# type: ...`
     pythonTypeNone          `None` inside a type expression
     pythonTypeEllipsis      `...` inside a type expression
@@ -184,6 +186,12 @@ Type Annotation Groups:~
     pythonDocstring         Module / function / class docstrings
     pythonClassVar          self / cls / mcs
 
+    (*) Not highlighted by default, like the rest of Python's punctuation.
+    The same applies to pythonParams (parentheses of a def) and
+    pythonDefColon (the colon ending a def/class header). Link them to
+    color them: >
+        hi link pythonTypeBracket Delimiter
+<
 Core Groups:~
 
     pythonStatement         Keywords (def, class, return, type, etc.)
@@ -275,6 +283,9 @@ Manual syntax resync:~
     - `# type: ignore` stays a comment
     - Not loaded under :syntax off / manual or Neovim Tree-sitter
     - Removed ftplugin/python.vim, which had no effect
+    - Brackets, commas and colons in annotations and def/class headers are
+      no longer linked to Delimiter/Operator, so they look like the same
+      punctuation elsewhere (the groups still exist to link yourself)
     - g:python_enhanced_highlight_all no longer overwrites the individual
       g:python_highlight_* variables
     - Tests (tests/run.sh) run in CI on Vim (Linux/macOS/Windows) and Neovim
diff --git a/syntax/python.vim b/syntax/python.vim
index e160007..6a77964 100644
--- a/syntax/python.vim
+++ b/syntax/python.vim
@@ -643,9 +643,7 @@ hi def link pythonComment           Comment
 hi def link pythonTodo              Todo
 hi def link pythonShebang           Comment
 hi def link pythonEncoding          Comment
-hi def link pythonDefColon          Delimiter
 hi def link pythonDefComment        pythonComment
-hi def link pythonParams            Delimiter
 
 hi def link pythonString            String
 hi def link pythonRawString         String
@@ -694,14 +692,11 @@ endif
 
 if s:type_annotations
   hi def link pythonReturnArrow     Operator
-  hi def link pythonTypeColon       Operator
   hi def link pythonTypingType      Type
   hi def link pythonPrimitiveType   Type
   hi def link pythonTypeName        Type
   hi def link pythonTypeDotted      Type
   hi def link pythonTypeUnion       Operator
-  hi def link pythonTypeBracket     Delimiter
-  hi def link pythonTypeComma       Delimiter
   hi def link pythonTypeNone        Constant
   hi def link pythonTypeEllipsis    Constant
   hi def link pythonTypeString      String
diff --git a/tests/test_loading.vim b/tests/test_loading.vim
index 903873b..bafe415 100644
--- a/tests/test_loading.vim
+++ b/tests/test_loading.vim
@@ -65,6 +65,12 @@ call s:Check(s:loads == 1, 'sourced once on FileType, got ' . s:loads)
 " Palette is opt-in: by default groups keep their standard links
 call s:Check(execute('highlight pythonBuiltin') =~# 'links to Function',
       \ 'palette off by default')
+" Punctuation in annotations looks like punctuation elsewhere
+for s:group in ['pythonTypeBracket', 'pythonTypeComma', 'pythonTypeColon',
+      \ 'pythonParams', 'pythonDefColon']
+  call s:Check(execute('highlight ' . s:group) !~# 'links to',
+        \ s:group . ' not linked')
+endfor
 
 " 2. $VIMRUNTIME ahead of plugin: Vim's file wins the runtime! race, so the
 "    plugin must force-load ours -- every time, not only the first time